a quote from dr. Honda
You have 3 fuel pumps in the boat. Starting from the engine to the tank...
1) In the Vapor tank, there is a high pressure pump. It supplies the EFI rail with 36 psi. There is a regulator that returns extra fuel back to the tank. This system is like a toilet tank. The high pressure pump only makes high pressure... and has nothing to do with getting fuel into that tank. And, like a toilet, there is a float, and a valve to regulate the level in the tank. Because of this... you need pressure on the hose to feed the tank.
2) The square diaphragm pump on the block. This is your main fuel pump. It's 100% mechanical. On the back side... there is 2 vacuum ports that are open to the inside of the engine. These ports push and pull the diaphragm, and in turn... with the check valves, pumps the fuel from the tank, to the vapor tank. So... when you put it back on... make sure that the gasket is on the right way, and that it is sealed... AND... that you don't accidentally plug the holes with sealant.
The beauty of this type of pump is... that when it's full, and the diaphragm can't move... it simply stops pumping. BUT... if the lines are low/empty... it can move A LOT of volume. So... it's low pressure, with variable volume... and doesn't need a regulator.
3) The rectangle electric pump: This is your "Lift Pump". It only operates while cranking the engine. If you can think of an old outboard engine... there was always a "Fuel Bulb" in the system. And... before starting it for the day... you would reach in the back, and physically pump the bulb to make sure there was fuel in the line. (The fuel would drain back to the tank overnight) In our boats... the same thing happens. But, they replaced the bulb with the electric pump. SO... while at cranking speed... the diaphragm pump is VERY inefficient, and you would have to crank FOREVER if the filter was dry. So, they installed the electric pump to quickly suck fuel from the tank, and into the system. FYI... the 175 Sportjet didn't have this pump... and it was a bastard to start cold.